Well, I can't make any guarantees obviously, but I would be very surprised if Spore didn't have at least decent dynamic AI. In many games, we have already seen the AI forming a coherent system that reacts to the player but can also function independently of the player.
With respect to affecting the ecosystem in creature or civilization stages, I think Will and the team want to create a coherent whole. I remember seeing somewhere that when you are able to see the whole planet, you will see the very shore where you first emerged from the primordial ooze. By the same token, you should be able to eat a species to extinction and see the ecosystem adapt around this change. Then when you reach the space stage, you'll control that same kind of ecosystem from above and see how it connects back to your own world and your own experiences. At the least, we know from the San Francisco apple store footage that you can eat a species to extinction in the creature stage. This is the kind of dynamic idea that's getting me really excited for Spore.